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In addition, environmental factors exhibited an indirect negative influence on denitrification by negatively influencing the abundance of the nosZ gene. The comprehensive effects of the environmental factors in overlying water on NH3 volatilization were greater than those in sediment. The NH4+-N content, pH of overlying water and sediment salinity were the main controlling factors for NH3 volatilization in saline soils. Biochar application effectively regulated the denitrification rate by changing the environmental factors and denitrifying functional gene a
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