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1 ± 0.06 ‰) and heavy Cu isotopes (Δ65Cusorbed-solution = +0.17 ± 0.06 ‰) during the day at high pH and the excretion of Zn light isotopes (‒0.4 less then Δ66Znexcreted-biofilm less then +0.14 ‰) and Cu heavy isotopes (Δ65Cuexcreted-biofilm = +0.7 ± 0.3 ‰) during the night at lower pH. We interpreted Zn and Cu diel cycles as a combination of a desorption of EPS-metal complexes and a small active efflux during the night with an adsorption and an incorporation via an active uptake during the day. The hysteresis of metal concentration in so
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